NAPC 2001
June 26 - July 1 2001 Berkeley, California
Abstracts, O - Pe
(5/17/01)
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PALEOBIOGEOGRAPHIC AND PALEOCLIMATIC IMPLICATIONS
OF THE CENOZOIC MOLLUSCAN RECORD IN THE NORTHWESTERN PACIFIC
OLEINIK, Anton E., Dept. of Geography and Geology, Florida Atlantic University,
Boca Raton, FL, USA
The high latitude area of the Kamchatka Peninsula contains the northernmost-known
record of the Cenozoic molluscan faunas in the North Pacific, with the
vast majority of fossil localities situated above 56° N.
Coastal outcrops stretching for over 400 kilometers along the shores of
the western and northeastern Kamchatka are composed of shallow-marine
fossiliferous rocks deposited in a suite of marginal basins existed around
the paleo-Kamchatka volcanic arc during the Cenozoic. Evolutionary patterns
of Cenozoic high-latitude molluscan assemblages of Kamchatka, ranging
in age from the Early Paleocene through Early Pleistocene were heavily
controlled by climatic changes. Globally warm intervals were characterized
by low latitudinal gradients and broad biogeographic provinces with high
percentages of cosmopolitan taxa with warm-water affinities. Close similarities
between Paleocene and Eocene shallow-water molluscan assemblages of Kamchatka
and the western Coast of North America points on the existence of the
shallow-marine migration route along the southern margin of the Beringia
during most of Paleogene. Climatic deterioration started in the late Eocene
and continued throughout Early Oligocene increased the amount of diversification
and ecological adaptations within "future cold water" elements
of the biota and the stepwise extinction or southward retreat of the warm
water elements. Neogene assemblages were heavily influenced by climatic
fluctuations, connected with the intensification of the paleo-Kuroshio
current and episodic warming of the high latitudes, peaked at 15.5 and
13.4 Ma. Penetration of warm waters in the high latitudes facilitated
several northward migration pulses of temperate and subtropical genera.
These climate-controlled migratory events together with the background
climatic cooling throughout the Neogene, resulted in evolution and diversification
of the endemic faunal elements that characterize the numerous modern molluscan
provinces of the northern circum-Pacific.
EVOLUTION OF ZOOPHYCOS IN THE VOCONTIAN
THROUGH (SOUTHEASTERN FRANCE) FROM LOWER JURASSIC TO UPPER CRETACEOUS
OLIVERO, Davide, Université Lyon 1, Villeurbanne, France
The complex trace fossil Zoophycos is recorded worldwide, from
Cambrian to Holocene. It is still an enigmatic ichnofossil. Traces related
to Zoophycos have a significant general evolution in their organization,
with an increasing complexity in the burrow system, and in their environmental
position, with a general shift from the shelf to deep sea.
Ichnologically homogeneous material has been studied in a relatively
restricted place: the Vocontian Through, in southeastern France. Good
outcrops allow the study of Zoophycos from Toarcian to Cenomanian.
The host sediments are usually marly limestones, occasionally sandstones.
Zoophycos from Lower Jurassic are restricted to shelf environments,
they form regular simple helicoidal whorls and the outlines are rarely
lobate. The width of the laminae never exceeds 4050 cm. In sandy
facies of the same age morphologies are quite confusing, the structures
are rough and the dimensions more limited. The Middle Jurassic sediments
are characterized by large and slightly lobate Zoophycos. The paleobathymetric
range varies from outer shelf to middle slope. Lower Cretaceous traces
characterize deeper environments (lower slope) and their dimensions increase
up to 150 cm, with slight lobes always present. Locally, a few little
specimens are restricted to dysoxic intervals in some Valanginian deposits.
Very large Zoophycos with well-developed lobes have only been
recorded in deep-sea marly sediments (Aptian to Cenomanian), but also
at the top of Barremian turbidites surrounded by basinal limestones without
Zoophycos. Curiously, all the observed specimens are constructed
upwards in the sediment, except the well lobate forms, which are constructed
downwards. These are limited to mid-Cretaceous deep-sea substrates.
As a conclusion, this study confirms the general shift to deeper environments
and the increasing complexity in the morphology of the burrow system.
But this morphology seems to be more influenced by the substrate rather
than by the bathymetry alone.
SURVIVING TAPHONOMIC RUSSIAN ROULETTE: HOW
THE ODDS OF FOSSIL PRESERVATION AFFECT TIME-AVERAGING
OLSZEWSKI, T.D., Dept. of Geological Sciences, Indiana University, Bloomington,
IN, USA
The distribution of shell ages in a marine deposit places a limit on
the temporal resolution of questions that can be addressed using fossil
material. In modern settings, shell ages can range over 100s to 1000s
of years despite extremely rapid rates of destruction. This broad range
is thought to be due to mixing of recent shells with sub-fossil material
from below the taphonomically destructive zone (TDZ) but above the depth
of final burial (DFB). In this study, mixing and destructive processes
have been modeled stochastically to determine the influence of TDZ thickness,
DFB (reworking zone), and long-term burial rate on shell age distributions.
To model reworking, all shells above a randomly determined depth are mixed.
Probability of reworking decreases from the sediment-water interface (SWI)
to the DFB. To model taphonomic loss, a random number generator is used
to determine which shells survive and which do not (i.e., playing Russian
roulette). Probability of shell destruction decreases with depth from
the SWI to the base of the TDZ. After mixing, the depths of all shells
increase in accordance with a specified burial rate. For a single model
run, these three steps are repeated until all shells have been either
destroyed or buried below the DFB. For a given set of parameters, the
model produces expected shell age distributions, which are generally lognormal.
Increasing the TDZ relative to the DFB does not influence the mode or
range of the shell age distribution. Increasing the DFB increases the
age and number of the oldest shells and shifts the mode toward older shells.
Increasing the burial rate can shift modal ages older or younger, but
invariably increases the proportion of very old shells. These preliminary
results suggest that time-averaging is more a function of reworking and
burial rate than the rate of taphonomic destruction, which is primarily
reflected in the number of shells entering the rock record.
ALBIAN BENTHIC COMMUNITY OF THE EL ABRA FORMATION
FROM EAST-CENTRAL MEXICO
OMAÑA, Lourdes, and Gloria Alencaster, Instituto de Geología,
Universidad Nacional Autónoma de México, México
Cretaceous carbonate deposits of the El Abra Formation are widely distributed
in the folded Sierra Madre Oriental. At El Madroño locality this
stratigraphic unit contains an abundant, diverse and excellently preserved
paleobiota, composed of rudist and other kind of bivalves, gastropods,
corals, echinoids, sponges, foraminifers and algae, and it is located
in the NE extreme of the State of Queretaro, in eastern-central Mexico.
The preservation of the rudist specimens, complete and in three dimensions,
permitted us to redescribe the coalcomanid caprinid genera Texicaprina,
Mexicaprina and Kimbleia, and to propose the new genera Guzzyella,
Müllerriedia and Jalpania, which probably constitute a
new subfamily. This caprinid rudist fauna is 100% endemic at the generic
level. It occupies the marginal facies of the platform. Among the monopleurid
rudists present are Monopleura marcida, M. pinguiscula, and two
new species of this genus. In the lagoonal facies of the platform, Toucasia
texana is almost the only species. The foraminiferal association is
well preserved but scarcely diverse. The main species are Dictyoconus
walnutensis, Coskinolinoides texanus, Buccicrenata subgoodlandensis, Cuneolina
sp., and a few miliolids as Spiroloculina and Moesiloculina.
Also present is the planktic genus Favusella. Among the calcareous
algae are Cayeuxia piae, Lithocodium aggregatum and Acicularia
elongata. The age assignment of the deposit is based on the benthic
foraminifer Dictyoconus walnutensis, which is considered a regional
marker of the middle Albian. The paleoenvironmental reconstruction, inferred
from the lithology and benthic assemblage, suggests a warm shallow water
platform with optimal living conditions, with abundant nutrients and highly
oxygenated water, which permitted the notable flourishing of the community.
MICROVERTEBRATES FROM A NEW ARIKAREEAN LOCAL FAUNA,
JOHN DAY FORMATION, OREGON, WITH BIOSTRATIGRAPHIC AND TAPHONOMIC IMPLICATIONS
PAJAK, Alois F., III, Theodore J. Fremd, and Dwight Hoy, John Day Fossil
Beds National Monument, Kimberly, OR, USA
Screen-washing techniques have generally been unproductive
within the late Oligocene and early Miocene strata of the John Day. As
a result, microvertebrates from the John Day Basin have largely been known
from isolated occurrences, with teeth and bones represented most frequently
by single elements. A newly discovered lithosympatric assemblage in the
northernmost exposures of the John Day Basin contains several Arikareean
age vertebrate faunas recovered from the previously unstudied Lone Rock
localities, which correlate to the upper John Day strata. Within this
assemblage, the Campbell Ranch Local Fauna consists of the largest microvertebrate
faunal concentration yet known from the John Day. The outcrop from which
the local fauna was recovered has been correlated with rocks in the Turtle
Cove area of the John Day Basin and consists of very fine silty tuffaceous
claystone in a loess-like deposit. This fauna represents a diverse population
of microvertebrates that, prior to the recovery of this material, were
not known to occur together within any stratigraphic intervals defined
in the John Day strata. Likewise, the paucity of microvertebrate occurrences
within the John Day has not permitted any detailed population or taphonomic
studies prior to the recovery of the over 5000 specimens recovered so
far from this locality. This local fauna is dominated by geomyids and
cricetids, with insectivores, leporids, heteromids, and aplodontids forming
about 15% of the remaining material. The first reported local occurrence
of Plesiosminthus sp. and other taxa within the John Day strata
assists in confirming a late Arikareean age for this fauna. The assemblage
is of taphonomic interest, representing a mode of accumulation different
from any previously identified localities in the John Day Basin. Preliminary
examination indicates the fossil remains are consistent with descriptions
of microvertebrate remains recovered from owl middens.
PLEISTOCENE PERSISTENCE AND THE RECENT DECLINE
IN CARIBBEAN CORAL REEF COMMUNITIES
PANDOLFI, John M., Dept. of Paleobiology, National Museum of Natural
History, Smithsonian Institution, Washington, DC, USA
The recent fossil history of coral reefs can provide
a critical window in which to view the magnitude, scale, and frequency
of change over significantly broader time intervals than available to
modern ecology. I examined the structure of Pleistocene Caribbean coral
communities using a hierarchical sampling design at broad spatial and
temporal scales. Significant differences in the composition of coral communities
from the leeward reef crest among three islands (San Andrés, Curaçao,
and Barbados) during the last interglacial, 125 ka (thousand years ago),
were driven by variability in the relative abundance of the same four
or five abundant taxa. At Barbados, coral composition remained constant
from 220-125 ka, but differed during the 104 ky reef-building episode.
However, the 104 ky community was closer in composition to older coral
communities from Barbados than it was to communities from San Andrés
or Curaçao. Separate analyses on the composition of the rare taxa
(data compiled using 1 hr searches) and those of the common taxa (data
compiled using 40 m transects) gave highly concordant results, suggesting
the composition of the rare taxa is correlated with that of the common,
structurally dominant corals.
These Pleistocene data point to a high degree of order
in coral communities over broad spatial and temporal scales and support
the importance of local influences in determining reef coral community
structure. Moreover, they indicate a structure to reef communities that
persisted to the 1980s, when rapid habitat degradation led to the collapse
or severe alteration of coral communities throughout the Caribbean. Whether
such habitat reduction and change in species composition is unprecedented
and due to human consumption and pollution, or it represents a short-term
fluctuation in an otherwise predictable community structure, is one of
the most important questions facing reef managers today. Present trends
are not predicted from history but may well be related to human-induced
environmental modification.
INVADING THE LAND AND SEA: PROGRESS TOWARDS UNDERSTANDING
MAJOR ECOLOGICAL TRANSITIONS IN TURTLES AND THE ORIGINS OF SOME MODERN CLADES
PARHAM, James Ford, Museum of Paleontology, University of California,
Berkeley, CA, USA
Although turtles are commonly used as an example of conservative evolution,
they are capable of rapid evolutionary change and exhibit a wide array
of phenotypes during their long and storied history. Like other clades
that have adopted successful, yet divergent, variants on the tetrapod
Bauplan, the evolution of turtles is highly constrained, but homoplasy
is rampant. Homoplastic morphologies make phylogenetic reconstruction
based solely on morphology extremely difficult. In the absence of unambiguous
synapomorphies, temporal and geographic data can provide important, independent
lines of evidence. These data and the homoplasy observed in the impressive
fossil history of turtles, combined with data from their ecologically
diverse living relatives, allow us to make inferences about patterns of
turtle evolution using multiple lines of evidence. The invasions of the
land and sea are good examples of major ecological transitions resulting
in parallel changes in plesiomorphic, generalized grades. A consideration
of all of the available data suggests that invasions of the land and sea
evolved multiple times and that some of the currently recognized clades
are polyphyletic grades.
EVOLUTION AND DIVERSITY IN STYLOPHORAN ECHINODERMS
PARSLEY, Ronald L., Dept. of Geology, Tulane University, New Orleans,
LA, USA
Recent establishment and/or restudy of approximately
20 stylophoran genera enables us to differentiate, by cladistic analysis,
the Class Stylophora into well-defined subclades and to see a clear evolutionary
picture. Of nearly 70 known genera, 49 were analyzed using 57 unweighted
binary and multistate characters in PeeWee/Nona. Genera not included are
based on incomplete or poorly preserved specimens; most can be placed,
with reasonable confidence, within the proper subclade. Stylophora are
uncommon "homalozoan" echinoderms that originate in the Cambrian
and range into the L. Devonian, with a single Lazarus genus persisting
until the L. Pennsylvanian. The earliest stylophorans are cornutes from
the M. Cambrian of Bohemia and the Western United States. Key pleisiomorphic
characters are; asymmetrical thecae, primitive aulacophore with stylocone,
a ventral strut composed of M1 and M5 plate elements, and commonly, complex
sutural pore respiratory structures (cothurnopores, lamellipores). The
major evolutionary center(s) for the cornutes is in basins marginal to
Gondwana especially in the Montagne Noir, France and the "Barrandian"
region of Bohemia in the U. Cambrian and L. Ordovician. Ankyroids evolved
from the cornutes in the U. Cambrian, probably in the Montagne Noir region
and radiated worldwide from there. Key pleisiomorphic characters are:
bilateral thecal outlines, advanced aulacophores with styloids, detached
M5 portion of the strut that is expressed as the ventral CS somatic plate,
and no respiratory pore structures. In the L.M. Ordovician major
ankyroid diversification occurred resulting in least six subclades ("cornutiforms,"
mitrocystitids, peltocystids-kirkocystids, yachalicystids, anomalocystitids-placocystitids
and anomalocystitids-allanicytids) in the same Gondwana region. In each
subclade thecal reduction by loss of posterior marginals/somatic plates
has occurred. SilurianDevonian occurrence and diversification is
concentrated in seas marginal to Gondwana, especially in the Australian
and South African regions.
DEFINING THE RANGE OF SUDDEN AND GRADUAL EXTINCTION
SCENARIOS COMPATIBLE WITH ANY (SIMPLE) STRATIGRAPHIC DISTRIBUTION OF FOSSILS
PAYNE, Jonathan L., Dept. of Earth and Planetary Science, Harvard University,
Cambridge, MA, USA
Sudden extinction events can appear gradual in the fossil
record due to the Signor-Lipps effect. Springer (1990) demonstrated that
for a group of taxa the confidence levels of classical confidence intervals
reaching a putative extinction horizon should be uniformly distributed
if there was a sudden extinction at that horizon. The distribution of
confidence levels can be tested against the uniform distribution using
a goodness-of-fit test (e.g., Kolmogorov-Smirnov). This approach assumes
the null hypothesis of sudden extinction unless it is rejected on statistical
grounds. Sudden and gradual extinction, however, should be viewed as equally
likely alter natives and there will always be gradual extinction scenarios
that produce distributions of confidence levels that are indistinguishable
from those expected in the case of a sudden extinction. For a range of
gradual extinction scenarios, I used simulations to determine which would
produce distributions indistinguishable from a sudden extinction.
The ability to distinguish between sudden and gradual
extinction depends upon two factors: (1) the duration of the gradual extinction
as a function of the average gap between fossil occurrences, and (2) the
number of taxa included. The greater the duration of the gradual extinction,
the less likely it is to be mistaken for a sudden extinction. The range
of gradual extinction scenarios that are compatible with a sudden extinction
scenario decreases as the number of taxa increases. The simulation results
eliminate the need to consider sudden extinction separately from gradual
extinction; it is merely the most rapid possible version of gradual extinction.
The results can be applied to all situations that meet the assumptions
necessary for the calculation of classical confidence intervals. The use
of classical confidence intervals is expanded by the results of the simulations
by providing a new approach for constraining the minimum rate of gradual
extinction based upon information in the fossil record.
PENNSYLVANIAN BRACHIOPOD ASSEMBLAGE ZONES IN THE
ELY LIMESTONE OF EAST CENTRAL NEVADA
PEREZ-HUERTA, Alberto, and Norman M. Savage, Dept. of Geological Sciences,
University of Oregon, Eugene, OR, USA
The upper part of the Ely Limestone, exposed in the Illipah
Quadrangle of East Central Nevada, includes several horizons of silicified
brachiopods that are commonly preserved as in situ communities.
The formation comprises a sequence of biomicitic limestones that range
in thickness from 138 to 200 m with a Morrowan-Atokan age. At leat two
major brachiopod assemblage zones can be recognized in different sampling.
The "Spiriferid Assemblage Zone" is dominated by the genera
Reticulariina and Punctospirifer with the main fauna composed
of Composita ?subtilita, Cleiothyridina aff. C. elegans, Hustedia rotunda,
Eomarginifera nuda, and Kozlowskia ?splendens. The "Productid
Assemblage Zone" is dominated by the genera ?Inflatia and
?Rugoclostus with the main fauna composed of Punctospirifer
gnomus, Punctospirifer transversus, Reticulariina campestris, and
Hustedia ?brentwoodensis. It has been found that the genera Beecheria
and Rhipidomella and the species Cleiothyridina orbicularis
are present in both assemblage zones. The proportions of genera and the
composition of the brachiopod assemblage zones varies laterally and vertically
with local environmental changes that are also reflected by changes in
bottom sediment and associated bryzoans and corals. These may reflect
fluctuations in sea level, climate, or tectonic disturbances. Ongoing
work is directed toward further clarifying the ecological factors controlling
the compositional shifts in the assemblage zones. These factors may include
fluctuations in sea level, which was pointed out as the main factor by
Moffet and Langenheim (1986). These authors recognized in general, the
same "spiriferid and productid" brachiopod assemblage zones
and included both, in their microfacies 2 and microfacies 1 respectively.
These microfacies distinctions were part of their sedimentological division
into five microfacies that define 13 complete cycles from relatively deep
to shallow-water deposition.
STRATIGRAPHIC AND PALEOECOLOGIC IMPLICATIONS
OF A NEW SPECIES OF OSTREID FROM THE LOWER EOCENE VIENTO FORMATION, LA
POPA BASIN, NORTHEAST MEXICO
PERRILLIAT, Maria del Carmen, and Francisco Vega, Instituto de Geología,
UNAM, Ciudad Universitaria, Coyoacan, México
Oyster banks are abundant in the intertidal-fluviatile
facies of the Viento Formation in La Popa basin, Nuevo Leon. The size
of the oysters is remarkable, as the mean height of the specimens is 200
mm. A new species is referred to Ostrea (Turkostrea) n. sp., based
on ornamentation and presence of chomata on the internal margins of the
valves. Similar ostreids have been reported from Tunisia, where Ostrea
strictiplicata major Locard is present in Eocene beds. This species
has a triangular shape and the radial ribs of the left valve are more
numerous and more closely spaced than the new species from Nuevo Leon,
which is more ovate. These same features differentiate the new species
from Ostrea arrosis Aldrich from the Eocene of Texas. Both species
attain a large size. However, Ostrea (Turkostrea) n. sp. seems
to be larger and thicker, which may suggest a tropical climate for this
part of northeast Mexico during the lower Eocene. Presence of Turritella
mortoni postmortoni Conrad associated with some of the ostreid banks
where we collected the new species confirms a lower Eocene age for the
Viento Formation. The Viento Formation is one of the two youngest stratigraphic
units of the La Popa basin from which no age has been defined until now.
The overlying Carroza Formation remains undated, but it is possible that
its age will not be younger than Eocene. Distribution of the facies and
outcrops of the Viento Formation suggests that during lower Eocene times
the La Popa basin represented a fluvial landscape, one with considerable
amounts of organic matter in the rivers, which prevailed in this region
during a regression. Some of the oyster shells were used as substratum
by bryozoan colonies, whose growth reached considerable thickness.
LARGE-SCALE HETEROGENEITY IN THE STRATIGRAPHIC
RECORD: A SIGNIFICANT SOURCE OF BIAS IN GLOBAL DIVERSITY ESTIMATES
PETERS, Shanan E., and Michael Foote, Dept. of the Geophysical Sciences,
University of Chicago, Chicago, IL, USA
Many features of global diversity compilations have proven
robust to continued sampling and taxonomic revision. Inherent biases in
the stratigraphic record may nevertheless substantially affect estimates
of global taxonomic diversity. We use a simple estimate of the amount
of marine sedimentary rock available for sampling: the number of formations
in the stratigraphic Lexicon of the United States Geological Survey. We
find this to be positively correlated with two independent estimates of
rock availability: global outcrop area derived from the Paleogeographic
Atlas Project (University of Chicago) database, and percent continental
flooding. Epoch-to-epoch changes in the number of formations are strongly
correlated with changes in sampled Phanerozoic marine diversity at the
genus level. We agree with previous workers in finding evidence of a diversity-area
effect that is substantially weaker than the effect of the amount of preserved
sedimentary rock. Our results suggest that much of the observed, short-term
variation in marine diversity may be an artifact of variation, in the
amount, of rock available for study.
Like the comparison between change in number of formations
and change in sampled diversity, which addresses short-term variation
in apparent diversity, the comparison between absolute values of these
quantities, which relates to longer-term patterns also shows a positive
correlation. Moreover, there is no clear temporal trend in the residuals
of the regression of sampled diversity on number of formations. This raises
the possibility that taxonomic diversity may not have increased steadily
since the early Paleozoic. Because of limitations in our data, however,
this question must remain open.
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