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Pat Holroyd
Museum Scientist (Vertebrate Collections)

Pat Holroyd

Email: pholroyd@berkeley.edu

Phone: (510) 642-3733

Web page: http://www.ucmp.berkeley.edu/people/pah/index.html

Her research: "My research program is centered around understanding the pattern and process of change in the continental biota during warm intervals in the past. This could help us evaluate how long-term climate change may alter the Earth's biota in the future."

Burning questions: "How does climate change affect diversity? Do mammals and reptiles respond differently to climate change? What aspects of the biology and ecology of different taxa make them respond differently to climate change?"

Fossils that particularly interest her: "Fossil mammals and reptiles (principally turtles) from the Late Cretaceous and early Paleogene (approximately 70-30 million years ago). For the past ten years, I have maintained an active field program in the Greater Green River Basin of southwestern Wyoming, where the rocks preserve fossil mammals and reptiles from approximately 53 to 50 million years in age."

Publications:

Head, J. J., G. F. Gunnell, P. A. Holroyd, J. H. Hutchison, and R. L. Ciochon. 2013. Giant lizards occupied herbivorous mammalian ecospace during the Paleogene greenhouse in Southeast Asia. Proceedings of the Royal Society B: Biological Sciences 280: 20130665.  Read it


Kaufman, J.A., G.H. Turner, P.A. Holroyd, F. Rovero, and A. Grossman. 2013. Brain Volume of the Newly-Discovered Species Rhynchocyon udzungwensis (Mammalia: Afrotheria: Macroscelidea: Implications for Encephalization in Sengis. PLoS ONE 8(3): e58667. doi:10.1371/journal.pone.0058667  Read it


Brinkman, D.B., P. A. Holroyd, and J. D. Gardner (eds.) 2012. Morphology and Evolution of Turtles. Springer Netherlands, Dordrecht. 596 pages.


Parham, J.F., P.C.J. Donoghue, C.J. Bell, T.D. Calway, J.J. Head, P.A. Holroyd, J.G. Inoue, R.B. Irmis, W.G. Joyce, D.T. Ksepka, J.S.L. Patané, N.D. Smith, J.E. Tarver, M. Van Tuinen, Z. Yang, K.D. Angielczyk, J. Greenwood, C.A. Hipsley, L. Jacobs, P.J. Makovicky, J. Müller, K.T. Smith, J.M. Theodor, R.C.M. Warnock, and M.J. Benton. 2012. Best practices for justifying fossil calibrations. Systematic Biology 61(2):346-359.   Read it


Wheatley, P.V., Hofman, A.E., Holroyd, P.A., Goodwin, M.B., and Brown, S.T. 2011. Calcium isotope fractionations in vertebrates from modern and fossil ecosystems. Geological Society of America, Annual Meeting, Minneapolis, MN  Read it


Archibald, J.D., W. A. Clemens, K. Padian, T. Rowe, N. Macleod, P. M. Barrett, A. Gale, P. Holroyd, H.-D. Sues, N. C. Arens, J. R. Horner, G. P. Wilson, M. B. Goodwin, C. A. Brochu, D. L. Lofgren, S. H. Hurlbert, J. H. Hartman, D. A. Eberth, P. B. Wignall, P. J. Currie, A. Weil, G. V. R. Prasad, L. Dingus, V. Courtillot, A. Milner, A. Milner, S. Bajpai, D. J. Ward, and A. Sahni. 2010. Cretaceous Extinctions: Multiple Causes. Science. 328: 973


Gunnell, G.F., P.D. Gingerich and P.A. Holroyd. 2010. Ptolemaiida. Pp. 83-88 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.


Holroyd, P.A. 2010. Macroscelidea. Pp. 89-98 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.


Holroyd, P.A. 2010. Tubulidentata. Pp. 107-112 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.


Holroyd, P.A., F. Lihoreau, G. F. Gunnell, and E. Miller. 2010. Anthracotheriidae. Pp. 843-852 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.


Dirks, W., R.L. Anemone, P.A. Holroyd, D.J. Reid, and P. Walton. 2009. Phylogeny, life history, and the timing of molar crown formation in two archaic ungulates, Meniscotherium and Phenacodus (Mammalia, 'Condylarthra'). Pp. 3-8 in T. Koppe, G. Meyer, and K.W. Alt (eds.), Comparative Dental Morphology. Frontiers of Oral Biology 13. Karger Press, Basel.


Goodwin, M.B., T. Deméré, P. Holroyd, R. Wilson, and S. Dowker. 2009. Unusual preservation of fossil baleen (Cetacea: Mysticeti) from the Miocene of California, USA: Comparative morphology and stable isotope evidence for seasonality and growth. Journal of Vertebrate Paleontology 29(3):107A.


Grossman, A. and P.A. Holroyd. 2009. Miosengi butleri, gen. et sp. nov., (Macroscelidea) from the Kalodirr Member, Lothidok Formation, early Miocene of Kenya. Journal of Vertebrate Paleontology 29(3):957-960.


Holroyd, P. A. 2009. Past records of Elephantulus and Macroscelides: geographic and taxonomic issues. Afrotherian Conservation 7:3-7.  Read it


Holroyd, P. A. and N.J. Stevens. 2009. Differentiation of Phiomys andrewsi from Lavocatomys aequatorialis (n. gen., n. sp.) (Rodentia: Thryonomyoidea) in the Oligo-Miocene interval on continental Africa. Journal of Vertebrate Paleontology. 29(4):1331-1334.  Read it


Stevens, N.J., P.A. Holroyd, E.M. Roberts, P.M. O'Connor, and M.D. Gottfried. 2009. Kahawamys mbeyaensis (n. gen., n. sp.) (Rodentia: Thryonomyoidea) from the Late Oligocene Rukwa Rift Basin, Tanzania. Journal of Vertebrate Paleontology 29(2):631-634.   Read it


Bourque, J., J. Hutchison, P.A. Holroyd, and J. Bloch. 2008. A new kinosternoid (Testudines: Dermatemydidae) from the Paleocene-Eocene boundary of the Bighorn Basin, Wyoming and its paleoclimatological implications. Journal of Vertebrate Paleontology 27(3 Suppl.):55A.


Clementz, M. A., P. A. Holroyd, and P. L. Koch. 2008. Identifying aquatic habits of herbivorous mammals through stable isotope analysis. Palaios 23(9):574-585.  Read it


Clementz, M. T. and P. A. Holroyd. 2008. From river horses to sea cows: using stable isotopes in tooth enamel to identify semiaquatic mammals in the fossil record. The 6th International Conference on Applications of Stable Isotope Techniques to Ecological Studies. Honolulu, HI August 25-29.


Clementz, M. T., and P. A. Holroyd. 2008. Isotopic modeling of the hippo ecomorph: identifying aquatic habits of herbivorous mammals through stable isotope analysis. The 6th International Conference on Applications of Stable Isotope Techniques to Ecological Studies. Honolulu, HI August 25-29.


Clementz, M. T., P. A. Holroyd, and P. L. Koch. 2008. Identifying aquatic habits of herbivorous mammals through stable isotope analysis. Palaios 23(9): 547-585


Dirks, W., R. L. Anemone, P. A. Holroyd, D. J. Reid, and P. Walton. 2008. Phylogeny, life history and the timing of crown formation in two archaic ungulates, Meniscotherium and Phenacodus (Mammalia, “Condylarthra”), 14th International Symposium on Dental Morphology, Greifswald, Germany, August 27-30, Abstracts with Programs, p.O8


Goodwin, M. B. and P. A. Holroyd. 2008. Fractionation in fossil bioapatite: phylogeny as a confounding factor. Journal of Vertebrate Paleontology 28(3):85A


Head, J.J., and P.A. Holroyd. 2008. Assembly and biogeography of North American Paleogene snake faunas based on an expanded fossil record. Journal of Vertebrate Paleontology 27(3 Suppl.):90A.


Holroyd, P.A. & Strait, S.G. 2008. New data on Loveina (Primates: Omomyidae) from the early Eocene Wasatch Formation and implications for washakiin relationships, pp. 243-257, in J.G. Fleagle and C.C. Gilbert, Elwyn Simons: A Search for Origins. Springer.


Holroyd, P.A. 2008. New data on dental eruption patterns in condylarths and afrotheres. Journal of Vertebrate Paleontology 27(3 Suppl.):93A


Smith, K. T., B.-A. S. Bhullar, and P. A. Holroyd. 2008. Earliest African record of the Varanus stem-clade (Squamata: Varanidae) from the earliest Oligocene of Egypt. Journal of Vertebrate Paleontology 28(3): 909-913.


Clementz, M., and P. Holroyd. 2007. Using stable isotope analysis ot test inferred semi-aquatic habits in fossil herbivores from the early Oligocene of Egypt. Journal of Vertebrate Paleontology 27(3):59A.


Goodwin, M.B., P.G. Grant, G. Bench, and P. Holroyd. 2007. Elemental composition and diagenetic alteration of dinosaur bone: Distinguishing micron-scale spatial and compositional heterogeneity using PIXE. Palaeogeography, Palaeoclimatology, Palaeoecology 253:458-476. doi:10.1016/j.palaeo.2007.06.017.  Read it


Holroyd, P., and G. Gunnell. 2007. New data on Anthracotheriidae (Artiodactyla) from the Paleogene of Egypt. Journal of Vertebrate Paleontology 27(3):90A.


Holroyd, P.A., T. Tsubamoto, N. Egi, R. L. Ciochon, M. Takai, Soe Thura Tun, Chit Sein, and G.F. Gunnell. 2006. A rhinocerotid perissodactyl from the late middle Eocene Pondaung Formation, Myanmar. Journal of Vertebrate Paleontology 26(2):491-494.  Read it


Egi, N., P.A. Holroyd, T. Tsubamoto, Aung Naing Soe, M. Takai, and R.L. Ciochon. 2005. Proviverrine hyaenodontids (Creodonta: Mammalia) from the Eocene of Myanmar and phylogenetic analysis of the proviverrines from the Para-Tethys area. Journal of Systematic Paleontology 3(4):337-358.  Read it


Head, J.J., Holroyd, P.A., Hutchison J.H., and R.L. Ciochon. 2005. First report of snakes (Serpentes) from the late middle Eocene Pondaung Formation, Myanmar. Journal of Vertebrate Paleontology 25(1):246-250.


Holroyd, P.A., and J.C. Mussell. 2005. Origin and relationships of the Macroscelidea and Tubulidentata. Pp. 71-83 in K.D. Rose and J.D. Archibald (eds.), The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades. Johns Hopkins University Press, Baltimore, Maryland.


Holroyd, P.A., J.F. Parham, and J.H. Hutchison. 2005. A reappraisal of some Paleogene turtles from the southeastern United States. Journal of Vertebrate Paleontology 25(4):979-982.


Stidham, T.A., P.A. Holroyd, G.F. Gunnell, R.L. Ciochon, T. Tsubamoto, N. Egi, and M. Takai. 2005. An ibis-like bird (Aves: cf. Threskiornithidae) from the late middle Eocene of Myanmar. Contributions from the University of Michigan Museum of Paleontology 31(7):179-184.  Read it