University of California Museum of Paleontology UCMP in the field See the world (and its fossils) with UCMP's field notes.
About UCMP People Blog Online Exhibits Public programs Education Collections Research
About UCMP : Contact UCMP

Mark Goodwin
Assistant Director (Collections and Research)

Mark Goodwin


Phone: (510) 643-9745

Web page:

His research: Mark studies how dinosaur bones change with growth and how fossil bone is preserved.

Burning questions: "What's the difference chemically and mineralogically between an unaltered modern bone and a fossil bone? What is the effect, if any, of the burial environment on the mineral and chemical properties, and how do you measure it?"

Sidelights: Mark is also researching the behavior and life history of pachycephalosaurs (dome-headed dinosaurs) and ceratopsids (horned-dinosaurs), particularly Triceratops.

His favorite thing about research: "The field of paleontology is much more integrative now, and working with colleagues from geochemistry, biology, and medicine, makes research both fun and challenging. Probing fossils with particle beams and X-rays, or slicing thin sections only 100's of microns thick, provides a wealth of new questions and projects. Plus I enjoy traveling to new places and meeting new people!"


Boatman, E.M., M.B. Goodwin, H.N. Holman, S. Fakra and M.H. Schweitzer. 2016. Characterization of Tyrannosaurus structural proteins using synchrotron X-ray and infrared micro-beam techniques. Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts, p. 99. Salt Lake City.

Evans, D.C., T.E. Williamson, M.B. Goodwin and M. Loewen. 2016. New Stegoceras-grade pachycephalosaurs (Ornithischia) reveal a Late Campanian radiation of dome-headed dinosaurs in southern Laramidia. Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts, p. 132. Salt Lake City.

Goodwin, M.B. and D.C. Evans. 2016. The early expression of squamosal horns and parietal ornamentation confirmed by new end-stage juvenile Pachycephalosaurus fossils from the Upper Cretaceous Hell Creek Formation, Montana. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1078343.   Read it

Goodwin, M.B., J.A. Nirody, T. Huynh, J.R. Horner, D.Y. Parkinson, R.C. Schott and D.C. Evans. 2016. Mapping and visualizing the complex internal anatomy of pachycephalosaur domes. Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts, p. 145. Salt Lake City.

Wosik, M., M.B. Goodwin and D.C. Evans. 2016. A nestling-sized skeleton of Edmontosaurus (Ornithischia, Hadrosauridae) with an analysis of ontogenetic limb allometry in relation to major gait shifts. Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts, p. 254. Salt Lake City.

Cllites, E.C., M.B. Goodwin, C.R. Marshall. 2014. Making the former USGS Menlo Park invertebrate collection digitally accessible through the University of California Museum of Paleontology. Geological Society of America Abstracts with Programs. Vol.46, No.6, p.701.  Read it

Goodwin, M.B. 2014. Fossilization as a preservational process: New insights on bioapatite in the fossil record. Geological Society of America Abstracts with Programs. Vol. 68, No. 6, p.26.  Read it

Goodwin, M.B. and J.R. Horner. 2014. Cranial morphology of a juvenile Triceratops skull from the Hell Creek Formation, McCone County Montana, with comments on the fossil record of ontogenetically younger skulls. Pp. 333-347 in G.P. Wilson, W.A. Clemens, J.R. Horner, J.H. Hartman, (eds.), Through the End of the Cretaceous in the Type Locality of the Hell Creek Formation in Montana and Adjacent Areas. Geological Society of America Special Paper 503.

Holroyd, P.A., J.H. Hutchison, M.B. Goodwin. 2014. Turtles of the Judith River Formation and comparisons among Late Cretaceous faunas in western North America. Geological Society of America Abstracts with Programs, Vol. 46, No. 5, p. 98. Notes: Rocky Mountain (66th Annual) and Cordilleran (110th Annual) Joint Meeting (19-21 May 2014), Bozeman, MT.  Read it

Horner, J., M. Goodwin, and V. Weaver. 2014. Nerve-like structures in the enigmatic domes of pachycephalosaurids. Society of Vertebrate Paleontology 74th Annual Meeting Program & Abstracts, p. 150. Berlin.   Read it

Irmis, R.B., M.B. Goodwin, and G.P. Wilson. 2014. Early Mesozoic vertebrate assemblages from Ethiopia and their significance for Gondwanan biogeography. 4th International Palaeontological Congress. Programme and abstracts Mendoza, Argentina.

Schweitzer, M.H., W. Zheng, T.P. Cleland, M.B. Goodwin, E. Boatman, E. Theil, M.A. Marcus, S.C. Fakra. 2014. A role for iron and oxygen chemistry in preserving soft tissues, cells and molecules from deep time. Proceedings of the Royal Society B. v. 281:1775, 2013doi: 10.1098/rspb.2013.2741. Notes: This is a Proc.R.Soc. B 22 January 2014 publication, though first published online 27 November 2013.  Read it

Weaver, V., M. Goodwin, J. Horner. 2014. Histology of pachycephalosaurid crania reveals evidence of multiple tissue types and variable origin of apical pathologies. Society of Vertebrate Paleontology 74th Annual Meeting Program & Abstracts, p. 150. Berlin.  Read it

Boatman, E.M., R. Gronsky, M.B. Goodwin, and R.O. Ritchie. 2013. Elucidating the nanoscale structure of dinosaur bone. Microscopy Today, v.21(5):34-39. Notes: Cambridge University Press publication.  Read it

Goodwin, M. B. and J. R. Horner, R. Schott, and D. Evans. 2013. New Data on Developmental Cranial Ontogeny in Pachycephalosaurs. Journal of Vertebrate Paleontology Program and Abstracts 73rd Annual Meeting of the Society of Vertebrate Paleontology, Los Angeles, CA, p. 135.

Goodwin, M.B., R. Irmis, G. P. Wilson, B. Atnafu. 2013. Non-marine Mesozoic ecosystems of northeastern Africa: New insights from the northwest plateau of Ethiopia. Geological Society of America Abstracts with Programs, Vol. 45, No. 7:187-2.   Read it

Goodwin, M.B., T.A. Demere, E.M. Boatman, M.A. Marcus, and S.C. Fakra. 2013. The paleohistology, mineralogy, and elemental chemistry of baleen from the upper Miocene Purisima Formation (Santa Cruz Co.), Northern California, USA. The Second International Symposium on Paleohistology v. 2:54, Bozeman, MT.

Scannella, J., D. Fowler, M. Goodwin, J. Horner. 2013. The clandestine role of heterochrony in ceratopsian evolution as revealed by juvenile Triceratops. Journal of Vertebrate Paleontology Program and Abstracts 73rd Annual Meeting, Los Angeles, p. 206. Notes: SVP Annual Meeting abstract

E. Boatman, R. Gronsky, M. Goodwin. 2012. Microscopy and analytical X-ray characterization of preserved hierarchical structure in dinosaur bone. Microscopy and Microanalysis Vol. 18, Suppl 2:45.

Fowler, D. W., J. B. Scannella, M. B. Goodwin, and J. R. Horner. 2012. How to eat a Triceratops: Large samples of toothmarks provides new insight into the feeding behavior of Tyrannosaurus. Journal of Vertebrate Paleontology Program and Abstracts 72nd Annual Meeting, Raleigh, North Carolina, p. 96.

Goodwin, M. 2012. The University as a Portal to Collection-based Research, Outreach, and Education. Proceedings of APRU Research Symposium on University Museums: Forming a University Museum Collection Network as the Core of Frontier Research. Kyoto, Japan. Sept. 11-14, 2012. pp. 27-30.

Renne, P.R. and M.B. Goodwin. 2012. Direct U-Pb dating of Cretaceous and Paleocene dinosaur bones, San Juan Basin, New Mexico: COMMENT. Geology 2012;40;e259. doi: 10.1130/G32521C.1  Read it

Scannella, J. B., D. W. Fowler, M. B. Goodwin, and J. R. Horner. 2012. Transitional Triceratops: Details of an ontogenetic sequence from the upper middle unit of the Hell Creek Formation, Montana. Journal of Vertebrate Paleontology Program and Abstracts 72nd Annual Meeting, Raleigh, North Carolina, p. 166.

Boatman, E., S. Fakra, M. Matthew, M. Schweitzer, M. Goodwin. 2011. Composition of Tyrannosaurus and Brachylophosaurus soft tissues at high spatial resolution: New insights into protein diagenesis. Program and Abstracts 71st Annual Meeting, Las Vegas, p. 73.

Goodwin, M., K. Stanton, J. Horner, S. Carlson. 2011. Oxygen isotopic variability and preservation in Tyrannosaurus rex, modern ratites and crocodylians: Revisiting the thermophysiology of T. rex using Δ18O. Journal of Vertebrate Paleontology 31(3, suppl.):117-118A.  Read it

Hall, L. and M. Goodwin. 2011. A diverse dinosaur tooth assemblage from the Upper Jurassic of Ethiopia: Implications for Gondwanan dinosaur biogeography. Journal of Vertebrate Paleontology. Program and Abstracts 71st Annual Meeting, Las Vegas, p. 121.

Horner, J. R., M. B. Goodwin, N. Myhrvold. 2011. Dinosaur census reveals abundant Tyrannosaurus and rare ontogenetic stages in the Upper Cretaceous Hell Creek Formation (Maastrichtian), Montana, USA. PLoS ONE 6(2): e16574. doi:10.1371/journal.pone.0016574  Read it

Schott RK, Evans DC, Goodwin MB, Horner JR, Brown CM, et al. (2011) Cranial Ontogeny in Stegoceras validum (Dinosauria: Pachycephalosauria): A Quantitative Model of Pachycephalosaur Dome Growth and Variation. PLoS ONE 6(6): e21092. doi:10.1371/journal.pone.0021092  Read it

Wheatley, P.V., Hofman, A.E., Holroyd, P.A., Goodwin, M.B., and Brown, S.T. 2011. Calcium isotope fractionations in vertebrates from modern and fossil ecosystems. Geological Society of America, Annual Meeting, Minneapolis, MN  Read it

Archibald, J.D., W. A. Clemens, K. Padian, T. Rowe, N. Macleod, P. M. Barrett, A. Gale, P. Holroyd, H.-D. Sues, N. C. Arens, J. R. Horner, G. P. Wilson, M. B. Goodwin, C. A. Brochu, D. L. Lofgren, S. H. Hurlbert, J. H. Hartman, D. A. Eberth, P. B. Wignall, P. J. Currie, A. Weil, G. V. R. Prasad, L. Dingus, V. Courtillot, A. Milner, A. Milner, S. Bajpai, D. J. Ward, and A. Sahni. 2010. Cretaceous Extinctions: Multiple Causes. Science. 328: 973

Goodwin, M.B. 2010. California's first dino! Dig Magazine. 12(6):14-17

Goodwin, M.B. 2010. Look at me now! Dig Magazine. 12(6):18-21.

Goodwin, M.B. 2010. Take a closer look: how Triceratops grew its horns and why! Dig Magazine. 12(6):8-12.

Goodwin, M.B. and J.R. Horner. 2010. Historical Collecting Bias and the Fossil Record of Triceratops in Montana. pp. 551-563 in M.J. Ryan, B.J. Chinnery-Allgeier, and D.A. Eberth (eds.), New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium. Indiana University Press.

Schott, R., D. Evans, M. B. Goodwin, C. Brown, N. Longrich. 2010. Cranial ontogeny in Stegoceras and the evolution of cranial doming and ornamentation in Pachycephalosauria (Dinosauria: Ornithischia). Journal of Vertebrate Paleontology. Program and Abstracts 70th Annual Meeting, Pittsburgh 159(A).

Goodwin, M.B., T. Deméré, P. Holroyd, R. Wilson, and S. Dowker. 2009. Unusual preservation of fossil baleen (Cetacea: Mysticeti) from the Miocene of California, USA: Comparative morphology and stable isotope evidence for seasonality and growth. Journal of Vertebrate Paleontology 29(3):107A.

Horner, J.R and M.B. Goodwin. 2009. Extreme Cranial Ontogeny in the Upper Cretaceous Dinosaur Pachycephalosaurus. PLoS ONE 4(10): e7626. doi:10.1371/journal.pone.0007626   Read it

Schott, R.K., D.C. Evans, T.E. Williamson, T.D. Carr, and M.B. Goodwin. 2009. The anatomy and systematics of Colepiocephale lambei (Dinosauria: Pachycephalosauridae). Journal of Vertebrate Paleontology 29(3):771-786.

Andersen, D.W., M.B. Goodwin, N.C. Shostak, C.M. Wentworth, and J.C. Tinsley. 2008. A newly discovered mammoth in San Jose, California, poses a stratigraphic puzzle. Geological Society of America Abstracts with Programs. 40(1):59.

Goodwin, M. B. and P. A. Holroyd. 2008. Fractionation in fossil bioapatite: phylogeny as a confounding factor. Journal of Vertebrate Paleontology 28(3):85A

Goodwin, M.B. and W.A. Clemens. Mesozoic continental vertebrates from the Northwestern Plateau, Ethiopia. International conference on paleoanthropology, paleontology, and archaeology in Ethiopia. Jan. 12-14, 2008. Addis Ababa, Ethiopia. Program with abstracts, pp. 18.

Horner, J.R. and M.B.Goodwin. 2008. Ontogeny of cranial epi-ossifications in Triceratops. Journal of Vertebrate Paleontology 28(1):134-144.

Schott, R., D. Evans, T. Williamson, T. Carr, and M. B. Goodwin. 2008. Anatomy and systematics of Colepiocephale lambei (Ornithischia Pachycephalosauridae) revised. Journal of Vertebrate Paleontology 28(3):138A.

Schweitzer, M.H., R. Avci, T. Collier, and M.B. Goodwin. 2008. Microscopic, chemical and molecular methods for examining fossil preservation. Comptes Rendu. 7:159-184.  Read it

Clemens, W.A., M.B. Goodwin, J.H. Hutchison, C.R. Schaff, C.B. Wood, and M.W. Colbert. 2007. First record of a Jurassic mammal (?"Peramura") from Ethiopia. Acta Palaeontologica Polonica 52(3):433-439.  Read it

Goodwin, M.B., P.G. Grant, G. Bench, and P. Holroyd. 2007. Elemental composition and diagenetic alteration of dinosaur bone: Distinguishing micron-scale spatial and compositional heterogeneity using PIXE. Palaeogeography, Palaeoclimatology, Palaeoecology 253:458-476. doi:10.1016/j.palaeo.2007.06.017.  Read it

Hall, L. and M. Goodwin. 2007. A preliminary analysis of dinosaur teeth from the Mugher Mudstone of Ethiopia. Journal of Vertebrate Paleontology 27(3):86A.

Horner, J., M.B. Goodwin, and H. Woodward. 2007. Synonomy consequences of dinosaur cranial ontogeny. Journal of Vertebrate Paleontology 27(3):92A.

Goodwin, M. B., W. A. Clemens, J. R. Horner, and K. Padian. 2006. The smallest known Triceratops skull: New observations on ceratopsid cranial anatomy and ontogeny. Journal of Vertebrate Paleontology 26(1):103-112.  Read it

Horner, J. R. and M. B. Goodwin. 2006. Major cranial changes during Triceratops ontogeny. Proceedings of the Royal Society Biological Sciences 273:2757-2761   Read it

Goodwin, M. B. 2005. What good was all the headgear? For decoration. Natural History Magazine 114(4):48-49.

Lonardelli, I., H-R. Wenk, L. Lutterotti, and M.B. Goodwin. 2005. Texture analysis from synchrotron diffraction with the Rietveld method: Dinosaur tendon and salmon scale. Journal of Synchrotron Radiation 12:354-360.